Journal of Theoretical Biology

The Journal of Theoretical Biology is the leading forum for theoretical perspectives that give insight into biological processes. It covers a very wide range of topics and is of interest to biologists in many areas of research, including: • Brain and Neuroscience • Cancer Growth and Treatment • Cell Biology • Developmental Biology • Ecology • Evolution • Immunology, • Infectious and non-infectious Diseases, • Mathematical, Computational, Biophysical and Statistical Modeling • Microbiology, Molecular Biology, and Biochemistry • Networks and Complex Systems • Physiology • Pharmacodynamics • Animal Behavior and Game Theory Acceptable papers are those that bear significant importance on the biology per se being presented, and not on the mathematical analysis. Papers that include some data or experimental material bearing on theory will be considered, including those that contain comparative study, statistical data analysis, mathematical proof, computer simulations, experiments, field observations, or even philosophical arguments, which are all methods to support or reject theoretical ideas. However, there should be a concerted effort to make papers intelligible to biologists in the chosen field.

Quartiles

The set of journals have been ranked according to their SJR and divided into four equal groups, four quartiles. Q1 (green) comprises the quarter of the journals with the highest values, Q2 (yellow) the second highest values, Q3 (orange) the third highest values and Q4 (red) the lowest values.

Category	Year	Quartile
Agricultural and Biological Sciences (miscellaneous)	1999	Q1
Agricultural and Biological Sciences (miscellaneous)	2000	Q1
Agricultural and Biological Sciences (miscellaneous)	2001	Q1
Agricultural and Biological Sciences (miscellaneous)	2002	Q1
Agricultural and Biological Sciences (miscellaneous)	2003	Q1
Agricultural and Biological Sciences (miscellaneous)	2004	Q1
Agricultural and Biological Sciences (miscellaneous)	2005	Q1
Agricultural and Biological Sciences (miscellaneous)	2006	Q1
Agricultural and Biological Sciences (miscellaneous)	2007	Q1
Agricultural and Biological Sciences (miscellaneous)	2008	Q1
Agricultural and Biological Sciences (miscellaneous)	2009	Q1
Agricultural and Biological Sciences (miscellaneous)	2010	Q1
Agricultural and Biological Sciences (miscellaneous)	2011	Q1
Agricultural and Biological Sciences (miscellaneous)	2012	Q1
Agricultural and Biological Sciences (miscellaneous)	2013	Q1
Agricultural and Biological Sciences (miscellaneous)	2014	Q1
Agricultural and Biological Sciences (miscellaneous)	2015	Q1
Agricultural and Biological Sciences (miscellaneous)	2016	Q1
Agricultural and Biological Sciences (miscellaneous)	2017	Q1
Agricultural and Biological Sciences (miscellaneous)	2018	Q1
Agricultural and Biological Sciences (miscellaneous)	2019	Q1
Agricultural and Biological Sciences (miscellaneous)	2020	Q1
Agricultural and Biological Sciences (miscellaneous)	2021	Q1
Agricultural and Biological Sciences (miscellaneous)	2022	Q1
Agricultural and Biological Sciences (miscellaneous)	2023	Q2
Applied Mathematics	1999	Q2
Applied Mathematics	2000	Q2
Applied Mathematics	2001	Q2
Applied Mathematics	2002	Q2
Applied Mathematics	2003	Q2
Applied Mathematics	2004	Q1
Applied Mathematics	2005	Q1
Applied Mathematics	2006	Q1
Applied Mathematics	2007	Q1
Applied Mathematics	2008	Q2
Applied Mathematics	2009	Q1
Applied Mathematics	2010	Q1
Applied Mathematics	2011	Q2
Applied Mathematics	2012	Q2
Applied Mathematics	2013	Q2
Applied Mathematics	2014	Q1
Applied Mathematics	2015	Q2
Applied Mathematics	2016	Q2
Applied Mathematics	2017	Q2
Applied Mathematics	2018	Q2
Applied Mathematics	2019	Q2
Applied Mathematics	2020	Q2
Applied Mathematics	2021	Q2
Applied Mathematics	2022	Q2
Applied Mathematics	2023	Q2
Biochemistry, Genetics and Molecular Biology (miscellaneous)	1999	Q1
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2000	Q1
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2001	Q1
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2002	Q1
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2003	Q1
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2004	Q1
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2005	Q1
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2006	Q1
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2007	Q1
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2008	Q2
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2009	Q1
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2010	Q1
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2011	Q2
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2012	Q1
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2013	Q1
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2014	Q1
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2015	Q1
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2016	Q2
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2017	Q2
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2018	Q2
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2019	Q2
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2020	Q2
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2021	Q2
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2022	Q2
Biochemistry, Genetics and Molecular Biology (miscellaneous)	2023	Q2
Immunology and Microbiology (miscellaneous)	1999	Q2
Immunology and Microbiology (miscellaneous)	2000	Q1
Immunology and Microbiology (miscellaneous)	2001	Q1
Immunology and Microbiology (miscellaneous)	2002	Q1
Immunology and Microbiology (miscellaneous)	2003	Q1
Immunology and Microbiology (miscellaneous)	2004	Q2
Immunology and Microbiology (miscellaneous)	2005	Q2
Immunology and Microbiology (miscellaneous)	2006	Q2
Immunology and Microbiology (miscellaneous)	2007	Q2
Immunology and Microbiology (miscellaneous)	2008	Q2
Immunology and Microbiology (miscellaneous)	2009	Q2
Immunology and Microbiology (miscellaneous)	2010	Q2
Immunology and Microbiology (miscellaneous)	2011	Q2
Immunology and Microbiology (miscellaneous)	2012	Q2
Immunology and Microbiology (miscellaneous)	2013	Q2
Immunology and Microbiology (miscellaneous)	2014	Q2
Immunology and Microbiology (miscellaneous)	2015	Q2
Immunology and Microbiology (miscellaneous)	2016	Q2
Immunology and Microbiology (miscellaneous)	2017	Q3
Immunology and Microbiology (miscellaneous)	2018	Q2
Immunology and Microbiology (miscellaneous)	2019	Q3
Immunology and Microbiology (miscellaneous)	2020	Q3
Immunology and Microbiology (miscellaneous)	2021	Q2
Immunology and Microbiology (miscellaneous)	2022	Q2
Immunology and Microbiology (miscellaneous)	2023	Q2
Medicine (miscellaneous)	1999	Q1
Medicine (miscellaneous)	2000	Q1
Medicine (miscellaneous)	2001	Q1
Medicine (miscellaneous)	2002	Q1
Medicine (miscellaneous)	2003	Q1
Medicine (miscellaneous)	2004	Q1
Medicine (miscellaneous)	2005	Q1
Medicine (miscellaneous)	2006	Q1
Medicine (miscellaneous)	2007	Q1
Medicine (miscellaneous)	2008	Q1
Medicine (miscellaneous)	2009	Q1
Medicine (miscellaneous)	2010	Q1
Medicine (miscellaneous)	2011	Q1
Medicine (miscellaneous)	2012	Q1
Medicine (miscellaneous)	2013	Q1
Medicine (miscellaneous)	2014	Q1
Medicine (miscellaneous)	2015	Q1
Medicine (miscellaneous)	2016	Q1
Medicine (miscellaneous)	2017	Q2
Medicine (miscellaneous)	2018	Q2
Medicine (miscellaneous)	2019	Q2
Medicine (miscellaneous)	2020	Q2
Medicine (miscellaneous)	2021	Q2
Medicine (miscellaneous)	2022	Q2
Medicine (miscellaneous)	2023	Q2
Modeling and Simulation	1999	Q1
Modeling and Simulation	2000	Q1
Modeling and Simulation	2001	Q2
Modeling and Simulation	2002	Q1
Modeling and Simulation	2003	Q1
Modeling and Simulation	2004	Q1
Modeling and Simulation	2005	Q1
Modeling and Simulation	2006	Q1
Modeling and Simulation	2007	Q1
Modeling and Simulation	2008	Q1
Modeling and Simulation	2009	Q1
Modeling and Simulation	2010	Q1
Modeling and Simulation	2011	Q1
Modeling and Simulation	2012	Q1
Modeling and Simulation	2013	Q1
Modeling and Simulation	2014	Q1
Modeling and Simulation	2015	Q1
Modeling and Simulation	2016	Q1
Modeling and Simulation	2017	Q2
Modeling and Simulation	2018	Q2
Modeling and Simulation	2019	Q2
Modeling and Simulation	2020	Q2
Modeling and Simulation	2021	Q2
Modeling and Simulation	2022	Q2
Modeling and Simulation	2023	Q2
Statistics and Probability	1999	Q2
Statistics and Probability	2000	Q2
Statistics and Probability	2001	Q2
Statistics and Probability	2002	Q2
Statistics and Probability	2003	Q2
Statistics and Probability	2004	Q2
Statistics and Probability	2005	Q2
Statistics and Probability	2006	Q2
Statistics and Probability	2007	Q2
Statistics and Probability	2008	Q2
Statistics and Probability	2009	Q2
Statistics and Probability	2010	Q2
Statistics and Probability	2011	Q2
Statistics and Probability	2012	Q2
Statistics and Probability	2013	Q2
Statistics and Probability	2014	Q2
Statistics and Probability	2015	Q2
Statistics and Probability	2016	Q2
Statistics and Probability	2017	Q2
Statistics and Probability	2018	Q2
Statistics and Probability	2019	Q2
Statistics and Probability	2020	Q2
Statistics and Probability	2021	Q2
Statistics and Probability	2022	Q2
Statistics and Probability	2023	Q2

SJR

The SJR is a size-independent prestige indicator that ranks journals by their 'average prestige per article'. It is based on the idea that 'all citations are not created equal'. SJR is a measure of scientific influence of journals that accounts for both the number of citations received by a journal and the importance or prestige of the journals where such citations come from It measures the scientific influence of the average article in a journal, it expresses how central to the global scientific discussion an average article of the journal is.

Year	SJR
1999	0.839
2000	0.943
2001	0.839
2002	0.948
2003	1.040
2004	1.050
2005	1.111
2006	1.132
2007	1.277
2008	0.885
2009	1.139
2010	1.081
2011	0.810
2012	0.947
2013	1.044
2014	1.076
2015	1.072
2016	0.934
2017	0.746
2018	0.711
2019	0.568
2020	0.657
2021	0.601
2022	0.566
2023	0.553

Total Documents

Evolution of the number of published documents. All types of documents are considered, including citable and non citable documents.

Year	Documents
1999	211
2000	249
2001	228
2002	242
2003	270
2004	287
2005	266
2006	419
2007	407
2008	410
2009	414
2010	471
2011	440
2012	439
2013	363
2014	435
2015	481
2016	445
2017	383
2018	415
2019	365
2020	267
2021	266
2022	207
2023	196

Citations per document

This indicator counts the number of citations received by documents from a journal and divides them by the total number of documents published in that journal. The chart shows the evolution of the average number of times documents published in a journal in the past two, three and four years have been cited in the current year. The two years line is equivalent to journal impact factor ™ (Thomson Reuters) metric.

Cites per document	Year	Value
Cites / Doc. (4 years)	1999	1.379
Cites / Doc. (4 years)	2000	1.596
Cites / Doc. (4 years)	2001	1.479
Cites / Doc. (4 years)	2002	1.681
Cites / Doc. (4 years)	2003	1.812
Cites / Doc. (4 years)	2004	2.000
Cites / Doc. (4 years)	2005	2.276
Cites / Doc. (4 years)	2006	2.532
Cites / Doc. (4 years)	2007	2.656
Cites / Doc. (4 years)	2008	2.799
Cites / Doc. (4 years)	2009	2.942
Cites / Doc. (4 years)	2010	2.839
Cites / Doc. (4 years)	2011	2.748
Cites / Doc. (4 years)	2012	2.801
Cites / Doc. (4 years)	2013	2.726
Cites / Doc. (4 years)	2014	2.545
Cites / Doc. (4 years)	2015	2.503
Cites / Doc. (4 years)	2016	2.381
Cites / Doc. (4 years)	2017	2.088
Cites / Doc. (4 years)	2018	2.122
Cites / Doc. (4 years)	2019	2.334
Cites / Doc. (4 years)	2020	2.459
Cites / Doc. (4 years)	2021	2.371
Cites / Doc. (4 years)	2022	2.222
Cites / Doc. (4 years)	2023	1.982
Cites / Doc. (3 years)	1999	1.379
Cites / Doc. (3 years)	2000	1.554
Cites / Doc. (3 years)	2001	1.477
Cites / Doc. (3 years)	2002	1.676
Cites / Doc. (3 years)	2003	1.791
Cites / Doc. (3 years)	2004	1.904
Cites / Doc. (3 years)	2005	2.200
Cites / Doc. (3 years)	2006	2.580
Cites / Doc. (3 years)	2007	2.644
Cites / Doc. (3 years)	2008	2.736
Cites / Doc. (3 years)	2009	2.922
Cites / Doc. (3 years)	2010	2.729
Cites / Doc. (3 years)	2011	2.668
Cites / Doc. (3 years)	2012	2.755
Cites / Doc. (3 years)	2013	2.662
Cites / Doc. (3 years)	2014	2.535
Cites / Doc. (3 years)	2015	2.454
Cites / Doc. (3 years)	2016	2.339
Cites / Doc. (3 years)	2017	2.056
Cites / Doc. (3 years)	2018	2.078
Cites / Doc. (3 years)	2019	2.405
Cites / Doc. (3 years)	2020	2.617
Cites / Doc. (3 years)	2021	2.410
Cites / Doc. (3 years)	2022	2.242
Cites / Doc. (3 years)	2023	1.974
Cites / Doc. (2 years)	1999	1.298
Cites / Doc. (2 years)	2000	1.550
Cites / Doc. (2 years)	2001	1.404
Cites / Doc. (2 years)	2002	1.639
Cites / Doc. (2 years)	2003	1.634
Cites / Doc. (2 years)	2004	1.760
Cites / Doc. (2 years)	2005	2.208
Cites / Doc. (2 years)	2006	2.524
Cites / Doc. (2 years)	2007	2.474
Cites / Doc. (2 years)	2008	2.717
Cites / Doc. (2 years)	2009	2.792
Cites / Doc. (2 years)	2010	2.581
Cites / Doc. (2 years)	2011	2.443
Cites / Doc. (2 years)	2012	2.606
Cites / Doc. (2 years)	2013	2.651
Cites / Doc. (2 years)	2014	2.382
Cites / Doc. (2 years)	2015	2.332
Cites / Doc. (2 years)	2016	2.266
Cites / Doc. (2 years)	2017	1.963
Cites / Doc. (2 years)	2018	1.988
Cites / Doc. (2 years)	2019	2.530
Cites / Doc. (2 years)	2020	2.674
Cites / Doc. (2 years)	2021	2.347
Cites / Doc. (2 years)	2022	2.092
Cites / Doc. (2 years)	2023	1.985

Total Cites

Self-Cites

Evolution of the total number of citations and journal's self-citations received by a journal's published documents during the three previous years.
Journal Self-citation is defined as the number of citation from a journal citing article to articles published by the same journal.

Cites	Year	Value
Self Cites	1999	142
Self Cites	2000	138
Self Cites	2001	109
Self Cites	2002	94
Self Cites	2003	124
Self Cites	2004	122
Self Cites	2005	133
Self Cites	2006	216
Self Cites	2007	250
Self Cites	2008	288
Self Cites	2009	368
Self Cites	2010	327
Self Cites	2011	330
Self Cites	2012	372
Self Cites	2013	268
Self Cites	2014	303
Self Cites	2015	278
Self Cites	2016	314
Self Cites	2017	193
Self Cites	2018	237
Self Cites	2019	261
Self Cites	2020	116
Self Cites	2021	83
Self Cites	2022	62
Self Cites	2023	57
Total Cites	1999	1045
Total Cites	2000	1147
Total Cites	2001	1053
Total Cites	2002	1153
Total Cites	2003	1288
Total Cites	2004	1409
Total Cites	2005	1758
Total Cites	2006	2123
Total Cites	2007	2570
Total Cites	2008	2988
Total Cites	2009	3612
Total Cites	2010	3359
Total Cites	2011	3455
Total Cites	2012	3651
Total Cites	2013	3594
Total Cites	2014	3148
Total Cites	2015	3035
Total Cites	2016	2991
Total Cites	2017	2798
Total Cites	2018	2720
Total Cites	2019	2990
Total Cites	2020	3044
Total Cites	2021	2523
Total Cites	2022	2013
Total Cites	2023	1461

External Cites per Doc

Cites per Doc

Evolution of the number of total citation per document and external citation per document (i.e. journal self-citations removed) received by a journal's published documents during the three previous years. External citations are calculated by subtracting the number of self-citations from the total number of citations received by the journal’s documents.

Cites	Year	Value
External Cites per document	1999	1.191
External Cites per document	2000	1.367
External Cites per document	2001	1.324
External Cites per document	2002	1.539
External Cites per document	2003	1.619
External Cites per document	2004	1.739
External Cites per document	2005	2.034
External Cites per document	2006	2.317
External Cites per document	2007	2.387
External Cites per document	2008	2.473
External Cites per document	2009	2.625
External Cites per document	2010	2.463
External Cites per document	2011	2.413
External Cites per document	2012	2.475
External Cites per document	2013	2.464
External Cites per document	2014	2.291
External Cites per document	2015	2.229
External Cites per document	2016	2.093
External Cites per document	2017	1.914
External Cites per document	2018	1.897
External Cites per document	2019	2.195
External Cites per document	2020	2.518
External Cites per document	2021	2.330
External Cites per document	2022	2.173
External Cites per document	2023	1.897
Cites per document	1999	1.379
Cites per document	2000	1.554
Cites per document	2001	1.477
Cites per document	2002	1.676
Cites per document	2003	1.791
Cites per document	2004	1.904
Cites per document	2005	2.200
Cites per document	2006	2.580
Cites per document	2007	2.644
Cites per document	2008	2.736
Cites per document	2009	2.922
Cites per document	2010	2.729
Cites per document	2011	2.668
Cites per document	2012	2.755
Cites per document	2013	2.662
Cites per document	2014	2.535
Cites per document	2015	2.454
Cites per document	2016	2.339
Cites per document	2017	2.056
Cites per document	2018	2.078
Cites per document	2019	2.405
Cites per document	2020	2.617
Cites per document	2021	2.410
Cites per document	2022	2.242
Cites per document	2023	1.974

% International Collaboration

International Collaboration accounts for the articles that have been produced by researchers from several countries. The chart shows the ratio of a journal's documents signed by researchers from more than one country; that is including more than one country address.

Year	International Collaboration
1999	18.01
2000	19.28
2001	21.49
2002	22.31
2003	24.44
2004	25.44
2005	25.19
2006	28.88
2007	29.98
2008	27.07
2009	27.54
2010	29.51
2011	32.05
2012	32.35
2013	33.88
2014	30.34
2015	29.11
2016	31.46
2017	31.33
2018	32.77
2019	36.71
2020	38.20
2021	36.84
2022	28.02
2023	28.06

Citable documents

Non-citable documents

Not every article in a journal is considered primary research and therefore "citable", this chart shows the ratio of a journal's articles including substantial research (research articles, conference papers and reviews) in three year windows vs. those documents other than research articles, reviews and conference papers.

Documents	Year	Value
Non-citable documents	1999	30
Non-citable documents	2000	30
Non-citable documents	2001	33
Non-citable documents	2002	25
Non-citable documents	2003	17
Non-citable documents	2004	20
Non-citable documents	2005	22
Non-citable documents	2006	24
Non-citable documents	2007	25
Non-citable documents	2008	34
Non-citable documents	2009	41
Non-citable documents	2010	36
Non-citable documents	2011	29
Non-citable documents	2012	27
Non-citable documents	2013	28
Non-citable documents	2014	34
Non-citable documents	2015	35
Non-citable documents	2016	39
Non-citable documents	2017	33
Non-citable documents	2018	27
Non-citable documents	2019	20
Non-citable documents	2020	20
Non-citable documents	2021	19
Non-citable documents	2022	17
Non-citable documents	2023	11
Citable documents	1999	728
Citable documents	2000	708
Citable documents	2001	680
Citable documents	2002	663
Citable documents	2003	702
Citable documents	2004	720
Citable documents	2005	777
Citable documents	2006	799
Citable documents	2007	947
Citable documents	2008	1058
Citable documents	2009	1195
Citable documents	2010	1195
Citable documents	2011	1266
Citable documents	2012	1298
Citable documents	2013	1322
Citable documents	2014	1208
Citable documents	2015	1202
Citable documents	2016	1240
Citable documents	2017	1328
Citable documents	2018	1282
Citable documents	2019	1223
Citable documents	2020	1143
Citable documents	2021	1028
Citable documents	2022	881
Citable documents	2023	729

Cited documents

Uncited documents

Ratio of a journal's items, grouped in three years windows, that have been cited at least once vs. those not cited during the following year.

Documents	Year	Value
Uncited documents	1999	299
Uncited documents	2000	269
Uncited documents	2001	278
Uncited documents	2002	237
Uncited documents	2003	243
Uncited documents	2004	251
Uncited documents	2005	221
Uncited documents	2006	208
Uncited documents	2007	262
Uncited documents	2008	278
Uncited documents	2009	287
Uncited documents	2010	307
Uncited documents	2011	350
Uncited documents	2012	337
Uncited documents	2013	362
Uncited documents	2014	290
Uncited documents	2015	322
Uncited documents	2016	340
Uncited documents	2017	430
Uncited documents	2018	391
Uncited documents	2019	338
Uncited documents	2020	324
Uncited documents	2021	270
Uncited documents	2022	270
Uncited documents	2023	242
Cited documents	1999	459
Cited documents	2000	469
Cited documents	2001	435
Cited documents	2002	451
Cited documents	2003	476
Cited documents	2004	489
Cited documents	2005	578
Cited documents	2006	615
Cited documents	2007	710
Cited documents	2008	814
Cited documents	2009	949
Cited documents	2010	924
Cited documents	2011	945
Cited documents	2012	988
Cited documents	2013	988
Cited documents	2014	952
Cited documents	2015	915
Cited documents	2016	939
Cited documents	2017	931
Cited documents	2018	918
Cited documents	2019	905
Cited documents	2020	839
Cited documents	2021	777
Cited documents	2022	628
Cited documents	2023	498

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